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Dr. Andrew Hafs Professor



  • West Virginia University, PhD, 2011
  • Arkansas Tech University, MS, 2007
  • University of Wisconsin at Stevens Point, BS, 2005


Fall Semesters
  • BIOL 3361 Limnology
  • BIOL 4545 Fisheries Management
  • BIOL 4898 Fisheries Research I
Spring Semesters
  • BIOL 1500 Diversity of Life
  • BIOL 4534 Ichthyology
  • BIOL 4899 Fisheries Research II
  • Sci 6350 Computer Applications in Statistics
Summer Semesters
  • BIOL 2339 Ethics of Fish and Wildlife Management

Research Interests

My research interests are focused on the interaction between animal populations and aquatic environments. Specifically, I enjoy researching the influence that anthropogenic factors have on movement patterns and population dynamics for species of interest. It is my hope that determining the influence of anthropogenic factors, as they relate to interaction between animals and their habitat, we will be better prepared to conserve and rehabilitate the aquatic environments studied.

Recent Work

Glade, K.C., A.J. Kennedy, B.J. Miller, B.D. Erb, A.L. Thompson, A.W. Hafs. 2023. Comparisons of walleye fecundity before, during, and after rehabilitation of the Red Lakes Fishery. Journal of Fish and Wildlife Management.

Peterson, S.J., K.C. Glade, A.W. Hafs, D.L. Guelda, and R.W. Koch. 2023. Effects of artificial zebra mussel druses on macroinvertebrate communities and benthic carbon accumulation. Northeastern Naturalist 30:347-367.

Orgon, T.J., A.W. Hafs, C.W. Isaacson, and S.E. Bowe. 2023. Spatial and temporal variability of mercury in Upper and Lower Red Lake walleye. Ecotoxicology 32:811-823.

Glade, K.C., B.R. Herwig, T.D. Ahrenstorff, J.R. Reed, and A.W. Hafs. 2023. Diet patterns and niche overlap of Muskellunge and co-occurring piscivores in Minnesota lakes. North American Journal of Fisheries Management 43:656-676.

Skolte, A., C.W. Schoenebeck, and A.W. Hafs. 2022. Effects of a shallow lake condition shift on habitat, zooplankton, and yellow perch dynamics. North American Journal of Fisheries Management 42:659-667.

Renik, K.M. and A.W. Hafs. 2020. Effect of beaver on brook trout habitat in North Shore, Lake Superior, streams. North American Journal of Fisheries Management 40:427–445.

Graham, J.D., A.W. Hafs, and A.J. Kennedy. 2019. Does a bioenergetics model accurately predict fish consumption by American white pelicans? A case study on walleyes in the Tamarac River, Minnesota. North American Journal of Fisheries Management 39:485-497.

Johnson-Brice, S.M., K.M. Renik, S.K. Windels, and A.W. Hafs. 2018. A review of beaver–salmonid relationships and history of management actions in the Western Great Lakes (USA) region. North American Journal of Fisheries Management 38:1203-1225.

Graham, J.D., A.W. Hafs, and A.J. Kennedy. 2017. Quantification of walleye spawning substrate in a northern Minnesota river using side-scan sonar. North American Journal of Fisheries Management 37:420-428.

Hafs, A.W. and K.J. Hartman. 2017. Seasonal changes in condition of Appalachian brook trout. North American Journal of Fisheries Management 37:196-206.

Studinski, J.M., A.W. Hafs, J.M. Niles, and K.J. Hartman. 2016. The effects of riparian disturbance on the condition and summer diets of age-0 brook trout (Salvelinus fontinalis) in three central Appalachian streams. Canadian Journal of Fisheries and Aquatic Sciences 74:980-986.

Hartman, K.J., F.J. Margraf, A.W. Hafs, and M.K. Cox. 2015. Bioelectrical impedance analysis: a new tool for assessing fish condition. Fisheries 40:590-600.

Hafs, A.W. and K.J. Hartman. 2015. Development of temperature corrections for bioelectrical impedance analysis models developed for age-0 and adult brook trout. Journal of Fish Biology 86:304-316.

Hafs, A.W. and K.J. Hartman. 2014. Developing bioelectrical impedance analysis methods for age-0 brook trout. Fisheries Ecology and Management 21:366-373.

Hafs, A.W., L.R. Harrison, R.M. Utz, and T. Dunne. 2014. Quantifying the role of woody debris in providing bioenergetically favorable habitat for juvenile salmon. Ecological Modelling 285:30-38.

Wilberding, M.C. and A.W. Hafs. 2013. Angler catch-per-unit-effort in restored and reference sections of the Merced River, California: a preliminary analysis. California Fish and Game 99:149-154.

Hafs, A.W., P.M. Mazik, P.B. Kenney, and J.T. Silverstein. 2012. Impact of carbon dioxide level, water velocity, strain, and feeding regimen on growth and fillet attributes of cultured rainbow trout (Oncorhynchus mykiss). Aquaculture 350-353:46-53.

Hafs, A.W. 2011. Bioelectrical impedance analysis methods for prediction of brook trout Salvelinus fontinalis percent dry weight. Dissertation. West Virginia University, Morgantown, WV.

Hafs, A.W. and K.J. Hartman. 2011. Influence of electrode type and location upon bioelectrical impedance analysis measurements of brook trout. Transactions of the American Fisheries Society 140:1290-1297.

Hafs, A.W., J.M. Niles, and K.J. Hartman. 2011. Efficiency of gastric lavage on age-0 brook trout and the influence on growth and survival. North American Journal of Fisheries Management 31:530-534.

Hafs, A.W., C.D. Horn, P.M. Mazik, and K.J. Hartman. 2010. Influences of acid mine drainage and thermal enrichment on stream fish reproduction and larval survival. Northeastern Naturalist 17:575:592.

Hafs, A.W., C.J. Gagen, and J.K. Whalen. 2010. Smallmouth bass summer habitat use, movement, and survival in response to low flow in the Illinois Bayou, Arkansas. North American Journal of Fisheries Management 30:604-612.

Hafs, A.W. 2007. Smallmouth bass survival, movement, and habitat use in response to seasonally discontinuous surface flow. Master’s thesis. Arkansas Tech University, Russellville, AR.

Undergraduate Research in BSU Journal of Student Research

Pribyl, M. 2023. Effect of initial infestation of zebra mussels on freshwater drum diets in Red Lake, MN.

Mott, I. 2023. Effects of velocity and substrate size on the formation of zebra mussel druses.

Kvam, M. 2023. Have rusty crayfish affected the diets of yellow perch?

Zwiers, R. 2023. A diet comparison of bluegill and bluegill x pumpkinseed hybrids.

Radaich, Q.L. 2022. The effects of diet on yellow perch morphology throughout Lake Bemidji.

Severin, J. 2022. Changes in the Minnesota northern pike population over the past thirty years.

Hauser, A. 2022. Analysis of hematocrit levels in walleye Sander vitreus from similar waterbodies.

Kehne, M. 2022. Zebra mussel length-weight relationships across various substrate types in Lake Bemidji.

Carr, M. 2021. Exploring the genetic variability of Zebra Mussels Dreisenna polymoprha.

Sorensen, J. 2021. Effectiveness of endothall treatment in reducing Potamogeton crispus in Lake Julia.

Oxley K. 2021. Abiotic factors effect on summertime fish sampling.

Gibson, M. 2021. Burbot post-spawn diet.

Beck, C. 2021. Comparing the condition of deformed and non-deformed Walleyes in Long Lake, MN.

Steinmetz, W. and C. Tembrock. 2021. Stress response of Largemouth and Smallmouth Bass: barbless hooks verses barbed hooks c & r.

Holland, J. 2021. Predicting total length of fish in Minnesota lakes using bony structure dimensions.

Laurich, J. 2021. Factors influencing aquatic invertebrate diversity in mine pit lakes along the Mesabi Iron Range.

Fink, K. 2021. Effects of salt runoff on aquatic macroinvertebrate colonies.

Taylor, A. 2021. Is there a relationship between aquatic vegetation and amphipod stocking mortality?

Daleiden, C. and S. Renner. 2020. Walleye Sander vitreus visceral fat accumulation.

Sea, B. 2020. Using Northern Pike and Yellow Perch to attempt a trophic cascade.

Collins, M.S. and A.M. Medina. 2019. Examining Black Bullhead Ameiurus melas age and growth in shallow Minnesota lakes.

Mendel, J. 2019. Goldeye Hidon alosoides calorimetry in Red Lake.

Tangren, C. 2019. Effects of Rainbow Trout on Brook Trout population size in tributaries to Lake Superior in Cook County, Minnesota.

Miller, B.J. and B.D. Erb. 2019. Changes in Walleye fecundity and egg size before, during, and after rehabilitation of the Red Lakes fishery.

Wang, M. 2018. Hunter harvest effect on the breeding bird population of Northern Pintail Anas acuta in North America.

Amundson, J. 2018. Age-0 Walleye growth rates following spiny water flea Bythotrephes longimanus invasion.

Coyle, C. 2018. Seasonal fluctuations of zooplankton, dissolved oxygen, and temperature in Lake Bemidji, Minnesota.

Morey, K. and E. Larson. 2018. Growth rate and hatch date of age-0 fishes on Leech Lake, Minnesota before Zebra Mussel establishment.

Weidenborner, H. and J. Janecek. 2018. Burbot activity in relation to light penetration on Lake Bemidji, MN.

Henry, R. 2017. Effect of abiotic factors on Bluegill size structure in nine central Minnesota lakes.

Knutson, C. 2017. Influence of time spent in captivity on aggressive behaviors of gray wolves Canis lupus.

Lamb, W. 2017. Minnesota fish taxonomic key.

Peterson, S.J. 2017. Burbot growth rates in a northern mesotrophic lake.

Robinson, T. 2017. Effects of catch-and-release angling on egg densities in Largemouth Bass nests.

Tuckett, D. 2017. Relationship between zooplankton and Walleye rearing pond success.

Varela, W. and Z. Buckley. 2017. Redd habitat selection for Brook Trout in the Necktie River Bemidji, MN.

Oswald, P. 2016. Survival of Walleye eggs in Upper Red Lake and the Tamarac River.

Evans, M. 2015. Using Wild I.D. as a reliable source for mark and recapture studies on Northern Pike (Esox lucius).

DuFault, D. 2015. Abiotic habitat preferences of age-0 Paddlefish in Lake Sakakawea, North Dakota during late summer.

Klingsheim, B.J. 2015. Influence of depth and retrieval speed on Yellow Perch barotrauma recovery time in winter.

Orgon, T. 2015. Juvenile Chinook Salmon diet in comparison to invertebrate drift in the Merced River, California.

Spurbeck, A. 2015. Effects of cooler northern waters on growth rates in Bluegill and Pumpkinseed Sunfish.

Stott, N. 2015. Spatial and temporal factors influencing Northern Pike fry density estimates.

Zentner, D. 2015. A genetic based approach to management and stocking of Muskellunge.

Zentner, D. 2015. Assessing the contributions of two stocked strains on a Minnesota-Wisconsin border water.

Carrow, R. 2014. The effects of angling pressure on Northern Pike size structure.

Longtin, K. 2014. Changing body condition in Walleye with reference to ideal sample size requirements.

Oldenburg, C. 2014. Winter survival strategies of benthic invertebrates in temporary habitats in a northern Minnesota lake.

Powers, E. 2014. Defining the physiochemical, biological and habitat requirements for the upper extant of Oncorhynchus mykiss distribution.

Swenson, B.C. 2014. Effects of river current inflow on dissolved oxygen and temperature of refuge zones for Cisco in Lake Bemidji.

VanderWeyst, D. 2014. The effect of barometric pressure on feeding activity of Yellow Perch.

Vaske, M. 2014. Use of cast nets and seine hauls to estimate abundance of age-0 Yellow Perch.

Skoog, M.L. 2013. Brown Trout spawning location and relationship to suitable winter habitat.

Wilson, A. 2013. White Bass assessment in regards to changes in water level on Lake Traverse.

Puttonen, E.A. 2013 Comparison of a length size limit of northern pike population on two lakes in north central MN.

Graduate Students

Collin Mitchell (In Progress-Started August 2023) Burbot work

Marissa Pribyl (In Progress-Started August 2023) Hg – Red Lakes Ecosystem

Georgina Leramo (In Progress-Started August 2023) Yellow perch work

Abigail Fountain (In Progress-Started February 2022) Daily movement rate of burbot

Kendra Fink (In Progress-Started January 2022) The influence of rusty crayfish on yellow perch diets and growth

Edward Carlson (August 2020-December 2022) Environmental influences on the growth of inland cisco populations in three Minnesota sentinel lakes

Tyler Robinson (February 2019-April 2022) An assessment of burbot life history in Minnesota with a focus on movement dynamics and spawning vulnerability

Tyler Orgon (August 2018-January 2022) Spatial and temporal variability of mercury in Upper and Lower Red Lake walleye

Alicia Skolte (August 2019-December 2021) Implications of condition shifts on shallow lakes

Kamden Glade (January 2019-December 2021) Feeding patterns and diet overlap of Muskellunge and co-occurring piscivores in Minnesota Lakes

Joseph Amundson (January 2019-October 2021) Assessing the dispersal and recruitment of stocked walleye fry in a northern Minnesota chain of lakes

Kathryn Renik (August 2016-August 2019) Effect of beaver on brook trout habitat in north shore, Lake Superior streams

Steve Hauschildt (January 2016-January 2019) Spatial and temporal variability in post-larval yellow perch density

Phil Oswald (August 2016-September 2018) Adult walleye abundance and subsequent fry production in the Tamarac River, MN

Ethan Karppinen (August 2014-September 2018) Growth of pelagic age-0 yellow perch in relation to hatch dates and water temperature in north-central Minnesota lakes

John Kempe (January 2016-May 2018) Population characteristics of a closed northern pike (Esox lucius) catch-and-release population in Minnesota

Emily Powers (August 2014-March 2017) Estimating seasonal variation in condition of yellow perch

Jake Graham (January 2014-April 2016) Assessment of American white pelican predation and reproductive success of walleye in the Tamarac River, MN